What I'm asking is, is the only AC connection being made by connecting the two routers over Media Bridge, or do you actually have devices connecting over 802.11ac?
If the issue issue is only while connecting over a media bridge, then it means the issue might specifically be with the media bridge feature rather than wireless itself.
Just trying to isolate the issue some more.
I think the answer to your question is Yes, media bridge.
A more detailed answer about the test conditions:
Server/Internet end:
Modem -> AC66u #1 (in Router mode) ---> Server connected via Ethernet
5GHz devices: AC66u #2 (Bridge)
2.4GHz devices: Phone (n), Laptop/Server (n)
Office end:
PC + VOIP ---> AC66u (in Media Bridge mode) connected via Ethernet
AC66u #2 Bridged to AC66u #1 via 5GHz link.
File transfer tests are done with just a simple Samba share from Server to PC across the bridge. Other devices, while active, are not using bandwidth.
Distance between AC66u devices is about 50 feet, interior through 5 layers of sheetrock, horizontal plane only.
I do have a laptop available with 5GHz 802.11n capability, but everything else is 802.11n 2.4GHz (aside from the inactive Kindle which is 802.11g).